Even the Owenses' most severe critics acknowledge that Mark's approach to conservation saved the lives of elephants.But the price, Markus Borner suggested, was too steep.

即便对欧文斯夫妇最严厉的批评者都承认马克·欧文斯的保护行动挽救了大象的生命，但马库斯·伯恩暗示代价过高了。

Even the Owenses' most severe critics acknowledge that Mark's approach to conservation saved the lives of elephants. But the price, Markus Borner suggested, was too steep.

即便对欧文斯夫妇最严厉的批评者都承认马克·欧文斯的保护行动挽救了大象的生命，但马库斯·伯恩暗示代价过高了。