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His misunderstanding of language was the primary cause of his other problems.
他对语言的误解是他其他问题的主要原因。
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You have to know the primary cause of it.
你必须知道的首要原因的。
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A primary cause of Tom's failure is his laziness.
懒惰是汤姆失败的主要原因之一。
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The primary cause of Tom's failure is his laziness.
汤姆失败的根本原因是懒惰。
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But female choice probably is not the primary cause of the traits.
但是,女性的选择或许不是导致这些特征的首要原因。